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Moldova is one of the smaller countries of the Danube River Basin, but the Danube touches the southern point of Moldova for about 340 meters. Picturesque Moldova holds over 12,500km² of the Danube River Basin, including 8,300km² of the Prut River Sub-Basin, 3,300km² of the Yalpugh River Sub-Basin and 900km² of the Cahul River Sub-Basin.

The Danube begins at the confluence of the Brigach and Breg Rivers in the city of Donaueschingen located in the Black Forest, Baden-Württemberg. The German Danube catchment area covers an area of about 56,200 km²; its share of the total Danube catchment area is approx. 7%. The German Danube basin is bordered by the Rhine basin to the west and the Elbe River basin to the east.

With a national territory of 87,609 km2, Croatia is at the intersection of the Pannonian Plain, the Balkans and the Adriatic. The country straddles the border of two major catchment areas: the Danube Basin and the Adriatic Sea. Draining over 62% of Croatia’s mainland, the Danube Basin covers the northern and central inland section of the country and is home to 69% of the population. Croatian territory accounts for 4.4% of the entire Danube Basin.

As both the largest remaining natural wetland and second largest river delta in Europe, the Danube Delta is one of the continent's most valuable habitats for wetland wildlife. Pollution and discharge manipulation from upstream has a huge effect on this area of high biodiversity, plus the delta continues to change ecologically itself – the Danube Delta spreads seaward at a rate of 10 to 25 metres every year.

The Sava is the Danube's largest tributary of the Danube in terms of discharge and the second largest in terms of catchment area. The Sava is shared by Slovenia, Croatia, Bosnia and Herzegovina, Serbia and Montenegro. The joint management arrangements acted as a pilot for the implementation of the European Union's Water Framework Directive for the Danube and Europe.

The Drava River is the fourth largest, as well as the fourth longest Danube tributary. It connects the Alps with the Danube and the Black Sea. The Drava has been considerably regulated with dams constructed to generate hydroelectricity and channels dredged to direct its flow. Nevertheless, natural habitats along the middle and lower reaches host unique varieties of flora and fauna, and several endemic species.